Any Windows or Linux client will have DNS query tools available. So connect one to your LAN and run the nslookup/dig command I mentioned earlier. Target an external DNS server in your query to see whether you get a response. If you can ping 8.8.8.8, for instance, but don't get a reply when running 'nslookup www.google.com 8.8.8.8' then I would look carefully at your firewall rules. If in doubt, post them and maybe someone can help further. Otherwise, I think we've just about exhausted all possibilities at this stage.
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